Paytan Biogeochemistry Lab

photos of sunsets

Publications are listed in approximate order from most recent to earliest. Linked items are PDF reprints. A complete list of Adina Paytan's publications, including those submitted and in press, can be found in her CV. Publication statistics can be found in Google Scholar.


Lecher, A.L., Murray, J. A. Paytan, (2018) Quantifying Processes Governing Nutrient Concentrations in a Coastal Aquifer via Principal Component Analysis. Hydrology 5, 15. DOI: 10.3390/hydrology5010015

Heck, N., A. Paytan, Potts, D.C., Haddad, B., Petersen, K. L. (2018) Management preferences and attitudes regarding environmental impacts from seawater desalination: Insights from a small coastal community. Elsevier Ocean and Coastal Management 163, Pp 22-29. DOI: 10.1016/j.ocecoaman.2018.05.024

Heck, N., Petersen, K. L., Potts, D.C., Haddad, B., Petersen, K. L. A. Paytan, (2018) Predictors of coastal stakeholders' knowledge about seawater desalination impacts on marine ecosystems. Elsevier Science of the Total Environmental 639 Pp. 785-792. DOI: 10.1016/j.scitotenv.2018.05.163

Martinez-Ruiz, F., Jroundi., A. Paytan, Guerra-Tschuschke, I., del Mar Abad, M. & González-Muñoz, M.T. (2018) Barium bioaccumulation by bacterial biofilms and implications for Ba cycling and use of Ba proxies. Nature Communications 9:1619 163, Pp 22-29. DOI: 10.1038/s41467-018-04069-z

Petersen, K.L., Frank, H., A. Paytan, Bar-Zeev, E. (2018) Impacts of Seawater Desalination on Coastal Environments. Chapter 11 in Sustainable Desalination Handbook: Plant Selection, Design and Implementation. Elsevier. Pp 437-463. DOI: 10.1016/B978-0-12-809240-8.00014-9

Haddad, B., Heck, N., A. Paytan, Potts, D. (2018) Social Issues and Public Acceptance of Seawater Desalination Plants. Chapter 9 in Sustainable Desalination Handbook: Plant Selection, Design and Implementation. Elsevier. Pp 505-525. DOI: 10.1016/B978-0-12-809240-8.00014-9

Slovak, N.M., Paytan, A., Rick, J.W., Chien, C. (2018) Establishing radiogenic strontium isotope signatures for Chavin de Huantar, Peru. Journal of Archaeological Science: Reports Volume 19, June 2018, Pages 411- 419, DOI: 10.1016/j.jasrep.2018.03.014

Reese, B.K., Zinke, L.A., Sobol, M.S., LaRowe, D.E., Orcutt, B.N., Zhang, X., Jaekel, U., Wang, F., Dittmar, T., Defforey, D., Tully, B., A. Paytan, Sylvan, J.B., Amend, J.P., Edwards K.J., Girguis, P. (2018): Nitrogen Cycling of Active Bacteria within Oligotrophic Sediment of the Mid-Atlantic Ridge Flank, Geomicrobiology Journal, DOI: 10.1080/01490451.2017.1392649

Silva-Tamayo, J.C., Lau, K.V., Jost, A.B., Payne, J.L., Wignall, P. B., Newton, R.J., Eisenhauer, A., Depaolo., D.J., Brown, S., Maher, K., Lehrmann, D.J., Altiner, D., Yu, M., Richoz, S., and A. Paytan (2018) Global perturbation of the marine calcium cycle during the Permian-Triassic transition. Geological Society of America, do:10.1130/B31818.1


Mellet, T., Selvin, C., Defforey, D., Roberts, K., Lecher, A.L., Dennis, K., Gutknecht, J., Field, C., A. Paytan (2017). Assessing Cumulative Effects of Climate Change Manipulations on Phosphorus Limitation in a Californian Grassland. Environmental Science and Technology. DOI: 10.1021/acs.est.7b04362.

Robinson, E., A.Paytan, Chien, C. (2017) Strontium isotope dates for the Oligocene Antigua Formation, Antigua, W. I. Caribbean Journal of Earth Science, 50, 11-18.;

Defforey, D. and A.Paytan (2017) Phosphorus cycling in marine sediments: Advances and Challenges. Chemical Geology 2017; doi:10.1016/j.chemgeo.2017.12.002

Lecher, A. I., Mackey, K. R. M., A.Paytan (2017) River and Submarine Groundwater Discharge Effects on Diatom Phytoplankton Abundance in the Gulf of Alaska. Hydrology 2017, 4, 61; doi:10.3390/hydrology4040061

Barbara Balestraa, B., Quintana Krupinski, N.B.,, Erohina T., Fessenden-Rahnc, J., Rahnc, T., A. Paytan (2017) Bottom-water oxygenation and environmental change in Santa Monica Basin, Southern California during the last 23 kyr. Palaeogeography, Palaeoclimatology, Palaeoecology. doi: 10.1016/j.palaeo.2017.09.002

Conrad, T.A., Nielsen, S.G., Peucker-Ehrenbrink, B., . Blusztajn, J., Winslow, D., Hein, J.R., A. Paytan (2017) Reconstructing the evolution of the submarine Monterey Canyon System from Os, Nd, and Pb isotopes in hydrogenetic Fe-Mn crusts doi: 10.1002/2017GC007071

Heck, N., A. Paytan, Potts, D. C., Haddad, B., Petersen, K.L. (2017) Management priorities for seawater desalination plants in a marine protected area: A multi-criteria analysis. Marine Policy 86, 64-71. doi: 10.1016/j.marpol.2017.09.12

Meija, L.M., Paytan, A., Eisenhauer, A., Bohm, F., Kolevica, A., Boltona, C., Mendez-Vicente, A., Abrevaya, L., Isensee, K., Stoll, H. (2018) Controls over 44/40Ca and Sr/Ca variations in coccoliths: New perspectives from laboratory cultures and cellular models (2017) Earth and Planetary Science Letters 481(2018) 48-60

Lau, K.V., Maher, K., Brown, S.T., Jost, A.B., Altiner, D., DePaolo, D. J., Eisenhauer, A., Kelley, B.M., Lehrmann, D.J., Paytan, A., Yu, M., Silva Tamayo, J.C., Payne, J.L. The influence of seawater carbonate chemistry, mineralogy, and diagenesis on calcium isotope variations in Lower-Middle Triassic carbonate rocks (2017) Chemical Geology doi: 10.1016/j.chemgeo.2017.09.006

A. Paytan (2017) The Inherent Conflict of Interest in For-Profit Journals, Eos Editor's Vox.

Fassbender, A.J., Palevsky, H.I., Martz, T.R., Ingalls, A.E., Gledhill, M., Fawcett, S.E., Brandes, J.A., Aluwihare, L.I., the participants of COME ABOARD, DISCO XXV. Perspectives on Chemical Oceanography in the 21st century: Participants of the COME ABOARD Meeting examine aspects of the field in the context of 40 years of DISCO. Marine Chemistry (2017), doi:10.1016/j.marchem.2017.09.002.

Paytan, A., Halverson, C., Weiss, E., Pedemonte, S., Mescioglu, E. (2017) Education for Climate Change Adaptation and Resilience, Limnology and Oceanography Bulletin.

Li, D., Zheng L.W, Jaccard, S.L., Fang, T.H., A. Paytan, Zheng, X., Chang Y.P. and Kao, S.J. (2017) Millennial-scale ocean dynamics controlled export productivity in the subtropical North Pacific, Geological Society of America, doi:10.1130/G38981.1.

Chuang, P.-C., Young, M.B., Dale, A.W., Miller, L.G., Herrera-Silveira, J.A. and A. Paytan (2017), A. Paytan. (2017) Methane fluxes from tropical coastal lagoons surrounded by mangroves, Yucatan, Mexico. Journal of Geophysical Research: Biogeosciences, 122, doi:10.1002/2017JG003761.

*Balestra, B., P. Grunert, B. Ausin, D. Hodell, J-A. Flores, C.A. Alvarez-Zarikian, F-J. Hernandez-Molina, D. Stow, W.E. Piller, A. Paytan (2017) Coccolithophore and benthic foraminifera distribution patterns in the Gulf of Cadiz and western Iberian margin during Integrated Ocean drilling program (IODP) expedition 339, 170, 50-67.

*Krupinski, N.B.Q., A.D. Russell, D.K. Pak, A. Paytan. (2017) Core-top calibration of B/Ca in Pacific Ocean Neogloboquadrina incompta and Globigerina bulloides as a surface water carbonate system proxy, Earth and Planetary Science Letters 466, 139-151.

Lecher, A.L., *P.C. Chuang, M. Singleton, A. Paytan. (2017) Sources of methane to an arctic lake in Alaska: An isotopic investigation, Journal of Geophysical Research: Biogeosciences, 122, doi:10.1002/2016JG003491/a>

Wang, F. J., *Y. Chen, Z. G. Guo, H. W. Gao, *K. R. Mackey, X. H. Yao, G. S. Zhuang, and , A. Paytan. (2017) Combined effects of iron and copper from atmospheric dry deposition on ocean productivity, Geophys. Res. Lett., 44, 2546 - 2555, doi:10.1002/2016GL072349.

*Defforey, D., B.J., Cade-Menun, A. Paytan. (2017) A new solution P NMR sample preparation scheme for marine sediments. Limnology and Oceanography: Methods, doi:10.1002/lom3.10166

Mackey, K.R.M., Kavanaugh, M.T., Wang, F., Chen, Y., Liu, F., Glover, D. M., Chien, C., A. Paytan (2017) Atmospheric and Fluvial Nutrients Fuel Algal Blooms in the East China Sea, Fronteirs in Marine Science 4, 2.

A. Paytan , Kathryn Roberts, Sue Watson, Sara Peek, Pei-Chuan Chuang, Delphine Defforey, Carol Kendall. (2017) Internal loading of phosphate in Lake Erie Central Basin, Science of the Total Environment 579, 1356-1365.

Chia-Te Chien, Tung-Yuan Ho, Matthew E. Sanborn, Qing-Zhu Yinc, and A. Paytan. (2017) Lead concentrations and isotopic compositions in the Western Philippine Sea, Marine Chemistry 189, 10-16.


Tracey Conrad, James R. Hein, A. Paytan and David A. Clague. (2016) Formation of Fe-Mn crusts within a continental margin environment, Ore Geol. Rev. (2016), http://

Peleg Astrahan, BarakHerut, A. Paytan and EyalRahav. (2016) The Impact of Dry Atmospheric Deposition on the Sea-Surface Microlayer in the SE Mediterranean Sea: An Experimental Approach. Front.Mar.Sci.3:222.doi: 10.3389/fmars.2016.00222

Alanna L. Lecher, Chia-Te Chien and A. Paytan. (2016) Submarine groundwater discharge as a source of nutrients to the North Pacific and Arctic coastal ocean. Marine Chemistry 186 (2016) 167–177.

Helen L. Cooper, Donald C. Potts and A. Paytan. (2016) Metabolic responses of the North Pacific krill, Euphausia pacifica, to short- and long-term pCO2 exposure. Mar Biol (2016) 163:207. DOI 10.1007/s00227-016-2982-z

Nadine Heck, A. Paytan, Donald C. Potts and Brent Haddad. (2016) Predictors of local support for a seawater desalination plant in a small coastal community. Environmental Science & Policy 66 (2016) 101–111.

Eyal Rahav, A. Paytan, Chia-Te Chien, Galit Ovadia, Timor Katz and Barak Herut. (2016) The Impact of Atmospheric Dry Deposition Associated Microbes on the Southeastern Mediterranean Sea Surface Water following an Intense Dust Storm. Front. Mar. Sci. 3:127. doi: 10.3389/fmars.2016.00127

Karen L. Knee, Elizabeth D. Crook, James L. Hench, James J. Leichter and A. Paytan. (2016) Assessment of Submarine Groundwater Discharge (SGD) as a Source of Dissolved Radium and Nutrients to Moorea (French Polynesia) Coastal Waters. Estuaries and Coasts, DOI 10.1007/s12237-016-0108-y

A. L Masterson, Boswell A. Wing, A. Paytan, James Farquhar and David T Johnston. (2016) The minor sulfur isotope composition of Cretaceous and Cenozoic seawater sulfate. Paleoceanography, doi:10.1002/2016PA002945

Paytan, A. and Bärbel Hönisch. (2016) Ocean Acidification - A Paleo Perspective. Limnology and Oceanography e-Lectures, doi:10.1002/loe2.10003.
e-Lecture link:

Pei-Chuan Chuang, Megan B. Young, Andrew W. Dale, Laurence G. Miller, Jorge A. Herrera-Silveira and A. Paytan. (2016) Methane and sulfate dynamics in sediments from mangrove-dominated tropical coastal lagoons, Yucatán, Mexico. Biogeosciences, 13, 2981–3001, doi:10.5194/bg-13-2981-2016

Chia-Te Chien, Katherine R. M. Mackey, Stephanie Dutkiewicz, Natalie M. Mahowald, Joseph M. Prospero and A. Paytan. (2016) Effects of African dust deposition on phytoplankton in the western tropical Atlantic Ocean off Barbados. Global Biogeochem. Cycles, 30, doi:10.1002/2015GB005334

Nadine Heck, A. Paytan, Donald Potts and Brent Haddad. (2016) Coastal residents' literacy about seawater desalination and its impacts on marine ecosystems in California. Marine Policy, 68, 178-186, doi:10.1093/icesjms/fsw021

Helen L. Cooper, Donald C. Potts, and A. Paytan. (2016) Effects of elevated pCO2 on the survival, growth, and moulting of the Pacific krill species, Euphausia pacifica
ICES J. Mar. Sci. doi:10.1093/icesjms/fsw021

Alistair J. Hobday, K. Cochrane, N. Downey-Breedt, J. Howard, S. Aswani, V. Byfield, G. Duggan, E. Duna, L. X. C. Dutra, S. D. Frusher, E. A. Fulton, L. Gammage, M. A. Gasalla, C. Griffiths, A. Guissamulo, M. Haward, A. Jarre, S. M. Jennings, T. Jordan, J. Joyner, N. K. Ramani, S. L. P. Shanmugasundaram, W. Malherbe, K. O. Cisneros, A. Paytan, G. T. Pecl, E´va E. Plaga´nyi, E. E. Popova, H. Razafindrainibe, M. Roberts, P. Rohit, S. S. Sainulabdeen, W. Sauer, S. T. Valappil, P. U. Zacharia and E. Ingrid van Putten. (2016) Planning adaptation to climate change in fast-warming marine regions with seafood-dependent coastal communities. Rev Fish Biol Fisheries. DOI 10.1007/s11160-016-9419-0

Alanna L. Lecher, Andrew T. Fisher and A. Paytan. (2016) Submarine groundwater discharge in Northern Monterey Bay, California: Evaluation by mixing and mass balance models. Marine Chemistry, 179, 44–55,

Elizabeth Derse Crook, Kristy J. Kroeker, Donald C. Potts, Mario Rebolledo-Vieyra, Laura M. Hernandez-Terrones and A. Paytan. (2016) Recruitment and Succession in a Tropical Benthic Community in Response to In-Situ Ocean Acidification. PLoS ONE 11(1): e0146707. doi:10.1371/journal.pone.0146707

Eyal Rahav, Galit Ovadia, A. Paytan and Barak Herut. (2016) Contribution of airborne microbes to bacterial production and N2 fixation in seawater upon aerosol deposition. Geophys. Res. Lett., 43, doi:10.1002/2015GL066898.

Stefan Markovic, A. Paytan, Hong Li and Ulrich G. Wortmann. (2016) A revised seawater sulfate oxygen isotope record for the last 4 Myr. Geochimica et Cosmochimica Acta 175 (2016) 239–251

Fenix Garcia-Tigreros Kodovsa, Katy J. Sparrow, Shari A. Yvon-Lewis, A. Paytan, Natasha T. Dimova, Alanna Lecher and John D. Kessler. (2016) Dissolved methane and carbon dioxide fluxes in Subarctic and Arctic regions: Assessing measurement techniques and spatial gradients. Earth and Planetary Science Letters, 436 (2016) 43–55


Zhongwu Ma, Christina Ravelo, Zhonghui Liu, Liping Zhou, and A. Paytan. (2015) Export Production Fluctuations in the Eastern Equatorial Pacific during the Pliocene-Pleistocene: Reconstruction Using Barite Accumulation Rates. Paleoceanography, 30, doi:10.1002/2015PA002860

Y. Zhang, N. Mahowald, R. A. Scanza, E. Journet, K. Desboeufs, S. Albani, J. F. Kok, G. Zhuang, Y. Chen, D. D. Cohen, A. Paytan, M. D. Patey, E. P. Achterberg, J. P. Engelbrecht, and K. W. Fomba (2015) Modeling the global emission, transport and deposition of trace elements associated with mineral dust. Biogeosciences, 12, 5771-5792, 2015, doi:10.5194/bg-12-5771-2015

Natasha T. Dimova, A. Paytan, John D. Kessler, Katy J. Sparrow, Fenix Garcia-Tigreros Kodovska, Alanna L. Lecher, Joseph Murray and Slawomir M. Tulaczyk (2015) Current Magnitude and Mechanisms of Groundwater Discharge in the Arctic: Case Study from Alaska. Environ. Sci. Technol, DOI: 10.1021/acs.est.5b02215

Andersson, A.J., D.I. Kline, P.J. Edmunds, S.D. Archer, N. Bednaršek, R.C. Carpenter, M . Chadsey, P. Goldstein, A.G. Grottoli, T.P. Hurst, A.L. King, J.E. Kubler, I.B. Kuffner, K.R.M. Mackey, B.A. Menge, A. Paytan, U. Riebesell, A. Schnetzer, M.E. Warner, and R.C. Zimmerman. (2015) Understanding ocean acidification impacts on organismal to ecological scales. Oceanography 28(2):16–27,

Delphine Defforey and A. Paytan (2015) Data report: characteristics of sedimentary phosphorusat North Pond, IODP Expedition 336. (2015) Proc. IODP | Volume 336 doi:10.2204/iodp.proc.336.205.2015

Alanna L. Lecher, John Kessler, Katy Sparrow, Fenix Garcia-Tigreros Kodovska, Natasha Dimova, Joseph Murray, Slawek Tulaczyk and A. Paytan (2015) Methane transport through submarine groundwater discharge to the North Pacific and Arctic Ocean at two Alaskan sites. (2015)

Alanna L. Lecher, Katherine Mackey, Raphael Kudela, John Ryan, Andrew Fisher, Joseph Murray and A. Paytan (2015) Nutrient Loading through Submarine Groundwater Discharge and Phytoplankton Growth in Monterey Bay, CA. Environmental Science & Technology. (2015)

Kathryn Roberts, Delphine Defforey, Benjamin L. Turner , Leo M. Condron, Sara Peek, Steve Silva, Carol Kendall and A. Paytan (2015) Oxygen isotopes of phosphate and soil phosphorus cycling across a 6500 year chronosequence under lowland temperate rainforest. Geoderma. (2015),

Laura M. Hernández-Terrones, Kimberly A.Null, Daniela Ortega-Camacho and A. Paytan (2015) Water quality assessment in the Mexican Caribbean : Impacts on the coastal ecosystem. Continental ShelfResearch. 102 (2015) 62–72,

S. Markovic, A. Paytan and U. G. Wortmann (2015) Pleistocene sediment offloading and the global sulfur cycle. Biogeosciences Discuss., 12, 1205–1245, 2015, doi:10.5194/bgd-12-1205-2015

Elizabeth M. Griffith, Matthew S. Fantle, Anton Eisenhauer, A. Paytan and Thomas D. Bullen (2015) Effects of ocean acidification on the marine calcium isotope record at the Paleocene–Eocene Thermal Maximum. Earth and Planetary Science Letters 419 (2015) 81–92,

Paytan, A., Alanna L. Lecher, Natasha Dimova, Katy J. Sparrow, Fenix Garcia-Tigreros Kodovska, Joseph Murray, Slawomir Tulaczyk, and John D. Kessler (2015) Methane transport from the active layer to lakes in the Arctic using Toolik Lake, Alaska, as a case study. Procedings of the National Academy of Science.

Mackey KRM, Chien C-T, Post AF, Saito MA and A. Paytan (2015) Rapid and gradual modes of aerosol trace metal dissolution in seawater. Front. Microbiol. 5:794. doi: 10.3389/fmicb.2014.00794


Iris Zohar, Barbara Cade-Menun, A. Paytan and Avi Shaviv (2014). Phosphorus Transformations from Reclaimed Wastewater to Irrigated Soil: A 31P NMR Study. Soil Sci. Soc. Am. J. 78:1884–1892 doi:10.2136/sssaj2014.01.0037.

C. Guieu, O. Aumont, A. Paytan, L. Bopp, C. S. Law, N. Mahowald, E. P. Achterberg, E. Marañón, B. Salihoglu, A. Crise, T. Wagener, B. Herut, K. Desboeufs, M. Kanakidou, N. Olgun, F. Peters, E. Pulido-Villena, A. Tovar-Sanchez, and C. Völker (2014). The significance of the episodic nature of atmospheric deposition to Low Nutrient Low Chlorophyll regions. Global Biogeochem. Cycles, 28, doi:10.1002/2014GB004852.

Mackey RM, Chia-Te Chein and A. Paytan (2014). Microbial and biogeochemical responses to projected future nitrate enrichment in the California upwelling system. Frontiers in Microbiology.

Jia Meng, Zhigang Yu, Qingzhen Yao, Thomas S. Bianchi, A. Paytan, Bin Zhao , Huihui Pan and Peng Yao (2014). Distribution, mixing behavior, and transformation of dissolved inorganic phosphorus and suspended particulate phosphorus along a salinity gradient in the Changjiang Estuary. Marine Chemistry.

Andrea M. Erhardt, Clare E. Reimers, David Kadko and A. Paytan (2014). Records of trace metals in sediments from the Oregon shelf and slope: Investigating the occurrence of hypoxia over the past several thousand years. Chemical Geology 382: 38-43.

Brian G. Burkhardt, Katie S. Watkins-Brandt, Delphine Defforey, A. Paytan and Angelicque E. White (2014). Remineralization of phytoplankton-derived organic matter by natural populations of heterotrophic bacteria. Marine Chemistry. doi: 10.1016/j.marchem.2014.03.007

Zhongwu Ma, Ellen Gray, Ellen Thomas, Brandon Murphy, James Zachos and A. Paytan (2014). Carbon sequestration during the Palaeocene–Eocene Thermal Maximum by an efficient biological pump. NATURE GEOSCIENCE | VOL 7 | MAY 2014. DOI: 10.1038/NGEO2139 Supplementary material 1 Supplementary material 2

Kimberly A. Null, Karen L. Knee, Elizabeth D. Crook, Nicholas R. de Sieyes, Mario Rebolledo-Vieyra, Laura Hernández-Terrones and A. Paytan (2014). Composition and fluxes of submarine groundwater along the Caribbean coast of the Yucatan Peninsula. Continental Shelf Research 77 (2014) 38–50.


Paytan, A, E. D. Crook, A. L. Cohen, T. R. Martz, Y. Takashita M. Rebolledo-Vieyra and L. Hernandez (2013). Reply to Iglesias-Prieto et al.: Combined field and laboratory approaches for the study of coral calcification. PNAS Early Edition.

K. McLaughlin, M. B. Young, A. Paytan, C. Kendall (2013). The O xygen Isotopic Compositi on of Phosphat e: a Tra cer for Phosphate Sources and Cycling. IAEA-TECDOC-1695

Mackey KRM, A. Paytan, K. Caldeira, A. R. Grossman, D. Moran, M. McIlvin, and M. A. Saito (2013). Effect of Temperature on Photosynthesis and Growth in Marine Synechococcus spp. Plant Physiology, Vol. 163, pp. 815–829, doi:

Crook, E.D., H. Cooper, D. C. Potts, T. Lambert and A. Paytan (2013). Food availability and pCO2 impacts on planulation, juvenile survival, and calcification of the azooxanthellate scleractinian coral, Balanophyllia elegans. Biogeosciences Discuss, 10, 7761–7783, 2013. doi:10.5194/bgd-10-7761-2013

Quintana Krupinski N.B., J R. Marlon, A Nishri, J H. Street and A. Paytan (2013). Climatic and human controls on the late Holocene fire history of northern Israel. Quaternary Research

Mackey KRM, D Hunter, E V. Fischer, Y Jiang, B Allen, Y Chen, A Liston, J Reuter, G Schladow and A. Paytan (2013). Aerosol-nutrient-induced picoplankton growth in Lake Tahoe. JOURNAL OF GEOPHYSICAL RESEARCH: BIOGEOSCIENCES, VOL. 118, 1–14, doi:10.1002/jgrg.20084, 2013

Crook, E.D., A. L. Cohen, M. Rebolledo-Vieyra, L. Hernandez and A. Paytan (2013). Reduced calcification and lack of acclimatization by coral colonies growing in areas of persistent natural acidification. Procedings of the National Academy of Science. 110(27), 11044-11049. doi/10.1073/pnas.1301589110.

McLaughlin, K., J. A. Sohm, G. A. Cutter, M. W. Lomas, and A. Paytan (2013). Phosphorus cycling in the Sargasso Sea: Investigation using the oxygen isotopic composition of phosphate, enzyme-labeled fluorescence, and turnover times. Global Biogeochem. Cycles, 27, 375-387. doi:10.1002/gbc.20037, 2013

J. Street, R. Anderson, R. Rosenbauer, and A. Paytan 2012. n-Alkane evidence for the onset of wetter conditions in the Sierra Nevada, California (USA) at the mid-late Holocene transition, ~3.0 ka. Quaternary Research 79:14-23 DOI:10.1016/j.yqres.2012.09.004

Erhardt AM, H. Pälike, and A. Paytan 2013. High-resolution record of export production in the eastern equatorial Pacific across the Eocene-Oligocene transition and relationships to global climatic records. Paleoceanography 2013

F. Jimenez-Espejo, J. Rodriguez-Vidal, C. Finlayson, F. Martinez-Ruiz, J. Carrion, A. Garcia-Alix, A. Paytan, F. Pacheco, D. Fa, G. Finlayson, M. Cortes-Sanchez, M. Gamiz, J. Gonzalez-Donoso, M. Linares, L. Caceres, S. Fernandez, K. Iijima, A. Aguirre 2012. Environmental conditions and geomorphologic changes during the Middle-Upper Palaeolithic in the southern Iberian Peninsula. Geomorphology DOI: 10.1016/j.geomorph.2012.10.011


F. Tamburini, S. M. Bernasconi and A. Paytan 2012. Phosphorus in the environment. Eos, Transactions American Geophysical Union. Vol. 93, Issue 41, page 405, 9 October 2012. DOI: 10.1029/2012EO410008

C. Buck and A. Paytan 2012. Evaluation of commonly used filter substrates for the measurement of aerosol trace element solubility. Limnol. Oceanogr.: Methods 10, 2012, 790–806. DOI: 10.4319/lom.2012.10.790

K Mackey, K Buck, J Casey. A Sid, M. Lomas, Y. Sohrin and A. Paytan 2012. Phytoplankton responses to atmospheric metal deposition in the coastal and open-ocean Sargasso Sea. Frontiers in Microbiology. 3(459) pp. 1-15. DOI: 10.3389/fmicb.2012.00359

A. Paytan and ET Gray. Sulphur Isotope Stratigraphy. Chapter 9 in "The Geologic Time Scale 2012" - 1st edition. Gradstein & Ogg & Schmitz & Ogg Editors, pp. 167-180. Elsevier. DOI: 10.1016/B978-0-444-59425-9.00009-3

Hinojosa, JL, ST Brown, J Chen, DJ DePaolo, A. Paytan, S Shen and JL Payne 2012. Evidence for end-Permian ocean acidification from calcium isotopes in biogenic apatite. Geology DOI: 10.1130/G33048.1

White, AE, KS Watkins-Brandt, MA Engle, B Burkhardt and A. Paytan 2012. Characterization of the rate and temperature sensitivities of bacterial remineralization of dissolved organic phosphorus compounds by natural populations. Frontiers in Microbiology Vol. 3 article 276. DOI: 10.3389/fmicb.2012.00276

Wortmann UG and A. Paytan 2012. Rapid Variability of Seawater Chemistry Over the Past 130 Million Years. Science 337, 334-346. DOI: 10.1126/science.1220656

Gonzalez-Muñoz MT, F Martinez-Ruiz, F Morcillo, JD Martin-Ramos , A. Paytan 2012. Precipitation of barite by marine bacteria: A possible mechanism for marine barite formation. Geology 40:675-678. DOI: 10.1130/G33006.1

Null KA, NT Dimova, KL Knee, BK Esser, PW Swarzenski, MJ Singleton, M Stacey, A. Paytan 2012. Submarine Groundwater Discharge-Derived Nutrient Loads to San Francisco Bay: Implications to Future Ecosystem Changes. Estuaries and Coasts. DOI: 10.1007/s12237-012-9526-7

Mackey KRM, K Roberts, MW Lomas, MA Sito, AF Post, A. Paytan, 2012. Enhanced Solubility and Ecological Impact of Atmospheric Phosphorus Deposition upon Extended Seawater Exposure. Env Sci & Tech Vol. 46:10438-10446. DOI: 10.1021/es3007996

Meeder E, K Mackey, A. Paytan, Y Shaked, D Iluz, N Stambler, T Rivlin, A Post, B Lazar 2012. Nitrite dynamics in the open ocean—clues from seasonal and diurnal variations. Mar Ecol Prog Ser Vol. 453: 11–26. DOI: 10.3354/meps09525

Street J, RS Anderson, and A. Paytan, 2012. An organic geochemical record of Sierra Nevada climate since the LGM from Swamp Lake, Yosemite. Quaternary Science Reviews 40: 89-106 DOI:10.1016/j.quascirev.2012.02.017

Griffith E and A. Paytan, 2012. Barite in the Ocean - Occurrence, Geochemistry, and Palaeoceanographic Applications. Sedimentology DOI: 10.1111/j.1365-3091.2012.01327.x

DeBond N, RL Oakes, A. Paytan, and UG Wortmann 2012. Early Aptian carbon and sulphur isotope signatures at ODP Site 765. Isotopes in Environmental and Health Studies. Vol. 48:1 180–194. DOI:10.1080/10256016.2012.659732

Crook ED, D Potts, M Rebolledo-Vierya, L Hernandez, A. Paytan, 2012. Calcifying coral abundance near low-pH springs: implications for future ocean acidification. Coral Reefs 31:239–245. DOI:10.1007/s00338-011-0839-y

Mackey K, C Mioni, J Ryan, and A. Paytan 2012. Phosphorus cycling in the red tide incubator region of Monterey Bay in response to upwelling. Frontiers in Microbiology. 3(33) DOI: 10.3389/fmicb.2012.00033

Paytan A, 2012. Mountains, Weathering and Climate. Science 335, 810; DOI: 10.1126/science.1218342

Sánchez M, F Espejo, M Vallejo, J Bao, A Carvalho, F Martinez-Ruiz, M Gamiz, J Flores, A. Paytan, J Lopez Sáez, L Peña-Chocarro, J Carrión, A Muñiz, Izquierdo, J Cantal, R Dean, E Salgueiro, R Martínez Sánchez, J De la Rubia de Gracia, M Lozano Francisco, J Vera Peláez, L Rodríguez, N Bicho. 2012. The Mesolithic–Neolithic transition in southern Iberia. Quat. Res., DOI:10.1016/ j.yqres.2011.12.003 (in press)

Knee, K and A. Paytan 2011. Submarine Groundwater Discharge: A Source of Nutrients, Metals, and Pollutants to the Coastal Ocean. In: Wolanski E and McLusky DS (eds.) Treatise on Estuarine and Coastal Science, Vol 4, pp. 205–233. DOI: 10 .1016/B978-0-12-374711-2 .00410-1


Hofmann GE, Smith, J., Johnson, K.S., Send, U., Levin, L.A., Micheli. F.,Paytan, A., Price, N.N., Peterson, B., Takeshita, Y., Matson, P.G., Crook, E. D., Kroeker, K.J., Gambi, M.C., Rivest, E.B., Frieder, C.A., Yu, P.C., Martz, T.R. 2011. High-Frequency Dynamics of Ocean pH: A Multi-Ecosystem Comparison. PLoS ONE 6(12): e28983. DOI:10.1371/journal.pone.0028983

Paytan, A, E T Gray, Z. Ma, A. Erhardt & K. Faul 2011. Application of Sulphur Isotopes for Stratigraphic Correlation. Isotopes in Environmental and Health Studies, 1-12. DOI:10.1080/10256016.2011.625423

Paytan, A and K. McLaughlin. 2011. Tracing the Sources and Biogeochemical Cycling of Phosphorus in Aquatic Systems Using Isotopes of Oxygen in Phosphate. M. Baskaran (ed.), Handbook of Environmental Isotope Geochemistry, Advances in Isotope Geochemistry, DOI 10.1007/978-3-642-10637-8_21

Slovak, NM and A. Paytan. 2011. Applications of Sr Isotopes in Archaeology. M. Baskaran (ed.), Handbook of Environmental Isotope Geochemistry, Advances in Isotope Geochemistry, DOI 10.1007/978-3-642-10637-8_35

Mackey, KRM, L. Bristow, D. Parks, M. Altabet, A. Post, and A. Paytan. 2011. The influence of light on nitrogen cycling and the primary nitrite maximum in a seasonally stratified sea. Progress in Oceanography, 545–560. DOI: 10.1016/j.pocean.2011.09.001

Knee, KL, E. Garcia-Solsona, J. Garcia-Orellana, A. Boehm, and A. Paytan. 2011. Using radium isotopes to characterize water ages and coastal mixing rates: A sensitivity analysis. Limnol. Oceanogr.: Methods, 380–395. DOI 10:4319/lom.2011.9.380

Chase, Z, A. Paytan, Beck, A., Biller, D., Bruland, K., Measurs, C., Sanudo-Wilhelmy, S. 2011. Evaluating the impact of atmospheric deposition on dissolved trace-metals in the Gulf of Aqaba, Red Sea, Mar. Chem. 126, 256-268. DOI:10.1016/j.marchem.2011.06.005

Griffith, EM, A. Paytan, A. Eisenhauer, D. Bullen, and E. Thomas. 2011. Seawater calcium isotope ratios across the Eocene-Oligocene transition. Geology 39(7): 683–686. DOI:10.1130/G31872.1

Post, AF, S. Penno, K. Zandbank, A. Paytan, S.M. Huse, and D.M. Welch. 2011. Long term seasonal dynamics of Synechococcus population structure in the Gulf of Aqaba, Northern Red Sea. Frontiers in Microbiology 2, DOI: 10.3389/fmicb.2011.00131

Gallego-Torres, D, F. Martinez-Ruiz, P. A. Meyers, A. Paytan, F. J. Jimenez-Espejo, and M. Ortega-Huertas. 2011. Productivity patterns and N-fixation associated with Pliocene-Holocene sapropels: paleoceanographic and paleoecological significance. Biogeosciences 8, 415–431

Wankel, SD, A. C. Mosier, C. M. Hansel, A. Paytan, and C. A. Francis. 2011. Spatial Variability in Nitrification Rates and Ammonia-Oxidizing Microbial Communities in the Agriculturally Impacted Elkhorn Slough Estuary, California. Applied and Environmental Microbiology 77(1): 269–280, DOI/10.1128/AEM.01318-10


Mackey, K. R. M., G. L. van Dijken, S. Mazloom, A. M. Erhardt, J. Ryan, K. R. Arrigo, and A. Paytan. 2010. Influence of atmospheric nutrients on primary productivity in a coastal upwelling region. Global Biogeochemical Cycles 24, GB4027, DOI/10.1029/2009GB003737.

de Sieyes, N. R., K. M. Yamahara, A. Paytan, and A. B. Boehm. 2010. Submarine groundwater discharge to a high-energy surf zone at Stinson Beach, California, estimated using radium isotopes. Estuaries and Coasts. DOI/10.1007/s12237-010-9305-2.

Feranec, R., E. A. Hadley and A. Paytan. 2010. Isotopes reveal limited effects of middle Pleistocene climate change on the ecology of mid-sized mammals. Quaternary International 217: 43-52.

Zohar, I., A. Shaviv, T. Klass, K. Roberts and A. Paytan. 2010. Method for the analysis of oxygen isotopic composition of soil phosphate fractions. Environmental Science and Technology 44: 7583-7588.

Zohar, I., Shaviv, A., Young, M., Kendall, C., Silva, S. and A. Paytan. 2010. Phosphorus dynamics in soils irrigated with reclaimed waste water or freshwater — A study using oxygen isotopic composition of phosphate. Geoderma 159: 109-121.

Knee, K.L., Gossett, R., Boehm, A.B., and A. Paytan. 2010. Caffeine and agricultural pesticide concentrations in surface water and groundwater on the north shore of Kauai (Hawaii, USA). Mar. Pollution Bulletin 60:1376-1382.

Cade-Menun, B.J and A. Paytan. 2010. Nutrient temperature and light stress alter phosphorus and carbon forms in culture-grown algae. Mar. Chem. DOI:10.1016/j.marchem.2010.03.002

Perry, E. C., G. Velazquez-Oliman, N. Wagner, A. Paytan and J. Street. 2010. Groundwater Sr, Cl, and SO4 chemistry in the western and southern Yucatan Peninsula: insight into formation of Edzna Valley, Mexico.

Griffith, E., M. Calhoun, E. Thomas, K. Averyt, A. Erhardt, T. Bralower, M. Lyle, A. Olivarez-Lyle, and A. Paytan. 2010. Export Productivity and Carbonate Accumulation in the Pacific Basin at the Transition from a Greenhouse to Icehouse Climate (Late Eocene to Early Oligocene). Paleooceanography 25, PA3212, DOI:10.1029/2010PA001932.

Payne, J. L, A.V. Turchynb, A. Paytan, D.J. DePaolod, D. J. Lehrmanne, M. Yuf, and J. Weig. 2010. Calcium isotope constraints on the end-Permian mass extinction. PNAS. Available online:

Landing, W. M. and A. Paytan. 2010. Aerosol chemistry and impacts on the ocean. Mar. Chem. 120: 1-3.

Knee, K.L., J.H. Street, E.E. Grossman, A.B. Boehm, and A. Paytan. 2010. Nutrient inputs to the coastal ocean from submarine groundwater discharge in a groundwater-dominated system: Relation to land use (Kona coast, Hawaii, U.S.A.). Limnol. Oceanogr. 55(3): 1105–1122.


Slovak, N.M. and A. Paytan. 2009. Fisherfolk and Farmers: Carbon and Nitrogen Isotope Evidence from Middle Horizon Anco´n, Peru. Int. J. Osteoarchaeol. DOI: 10.1002/oa.1128

Breier, J.A., N. Nidzieko, S. Monismith, W. Moore, and A. Paytan. 2009. Tidally regulated chemical fluxes across the sediment–water interface in Elkhorn Slough, California: Evidence from a coupled geochemical and hydrodynamic approach. Limnol. Oceanogr. 54(6), 2009, 1964–1980.

Young, M.B., K. McLaughlin, C. Kendall, W. Stringfellow, M. Rollog, K. Elsbury, E. Donald, and A. Paytan. 2009. Characterizing the Oxygen Isotopic Composition of Phosphate Sources to Aquatic Ecosystems. Environmental Science & Technology 43 (14): 5190-5196.

Black, F. J., A. Paytan, K. L. Knee, N. De Sieyes, P. Ganguli, E. Gray, A. R. Felgal. 2009. Submarine Groundwater Discharge of Total Mercury and Monomethylmercury to Central California Coastal Waters. Environmental Science & Technology 43 (15): 5652-5659.

Mackey, K. R. M. & A. Paytan. 2009. Phosphorus Cycle. Encyclopedia of Microbiology. Moselio Schaechter, Editor, pp. 322-334 Oxford: Elsevier.

Anderson, L. D., K.L. Faul, and A. Paytan. 2009. Phosphorus associations in aerosols: What can they tell us about P bioavailability? Mar. Chem. 120 (2010) 44–56.

Mackey, K. R. M, T. Rivlin, A.R. Grossman, A.F. Post, A. Paytan. 2009. Picophytoplankton responses to changing nutrient and light regimes during a bloom. Marine Biology doi/10.1007/s00227-009-1185-2.

Elsbury, K. E., A. Paytan, N.E. Ostrom, C. Kendall, M.B. Young, K. McLaughlin, M.E. Rollog, and S. Watson. 2009. Using Oxygen Isotopes of Phosphate To Trace Phosphorus Sources and Cycling in Lake Erie. Environmental Science & Technology 43: 3108–3114.

Paytan, A., K. R. M. Mackey, Y. Chen, I.D. Limac, S.C. Doneyc, N. Mahowaldd, R. Labiosae, and A.F. Postf. 2009. Toxicity of atmospheric aerosols on marine phytoplankton. Procedings from the National Academy of Science 106(12): 4601-4605. doi/10.1073/pnas.0811486106

Wankel, S., Y. Chen, C. Kendall, A.F. Post, A. Paytan. 2009. Sources of aerosol nitrate to the Gulf of Aqaba: Evidence from d15N and d18O of nitrate and trace metal chemistry. Mar. Chem. DOI:10.1016/j.marchem.2009.01.013.

Wankel, S. D., C. Kendall, and A. Paytan. 2009. Using nitrate dual isotopic composition (d15N and d18O) as a tool for exploring sources and cycling of nitrate in an estuarine system: Elkhorn Slough, California. Journal of Geophysical Research 114: G01011. DOI:10.1029/2008JG000729.

Perry, E., A. Paytan, B. Pedersen, G. Velazquez-Oliman. 2009. Groundwater geochemistry of the Yucatan Peninsula, Mexico: Constraints on stratigraphy and hydrogeology. Journal of Hydrology 367(1-2):27-40.

Foster,R. A., A. Paytan, and J. P. Zehr. 2009. Seasonality of N2 fixation and nifH gene diversity in the Gulf of Aqaba (Red Sea). Limnology and Oceanography 54(1): 219–233.

Raanan, H., A. Vengosh, A. Paytan, A. Nishri, A. Kabala. 2009. Quantifying saline groundwater flow into a freshwater lake using the Ra isotope quartet: A case study from the Sea of Galilee (Lake Kinneret), Israel. Limnology and Oceanography 54(1): 119–131.

Slovak, N. M. A. Paytan, B.A. Wiegnad. 2009. Reconstructing Middle Horizon mobility patterns on the coast of Peru through strontium isotope analysis. Journal of Archaeological Science 36: 157–165.


Griffith, E. M., A. Paytan, K. Caldeira, T.D. Bullem, E. Thomas. 2008. A Dynamic Marine Calcium Cycle During the Past 28 Million Years. Science 322: 1671. DOI: 10.1126/science.1163614

Grossman, E., J. B. Logan, J.H. Street, A. Paytan, P.S. Chavez, Jr. 2008. Ground water and its influence on reef evolution, Chapter 13 of Field, M.E., Cochran, S.A., Logan, J.B., and Storlazzi C.D., eds., The coral reef of south Moloka`i, Hawai`i; portrait of a sediment-threatened fringing reef: U.S. Geological Survey Scientific Investigations Report 2007-5101, p. 111-116 [].

Paytan, A. 2008. Ocean Paleoproductivity. In: Encyclopedia of Paleoclimatology and Ancient Environments. Vivien Gornitz, (Ed) Encyclopedia of Earth Science Series. Kluwer Academic Publishers.

Feranec, R.S., A.H. Hadley, A. Paytan. 2008. Stable isotopes reveal seasonal competition for resources between late Pleistocene bison (Bison) and horse (Equus) from Rancho La Brea, southern California. Palaeogeography, Palaeoclimatology, Palaeoecology 271:153–160.

Mahowald, N.M., S. Englestaedter, C. Luo, A. Sealy, P. Artaxo, C. Benitez-Nelson, S. Bonnet, Y. Chen, P.Y. Chuang, D.D. Cohen, F. Dulac, B. Herut, A.M. Johansen, N. Kubilay, R. Losno, W. Maenhaut, A. Paytan, J.M. Prospero, L.M. Shank, R.L. Siefert. 2008. Atmospheric Iron Deposition: Global Distribution, Variability, and Human Perturbations. Annual Review of Marine Science 1:245-78.

Griffith, E.M., E.A. Shauble, T.D. Bullen, A. Paytan. 2008. Characterization of calcium isotopes in natural and synthetic barite. Geochimica et Cosmochimica Acta. 72: 5641-5658

Knee, K.L., J. Stree, E. Grossman, A. Paytan. 2008. Submarine Ground-Water Discharge and Fate Along the Coast of Kaloko-Honokohau National Historical Park, Island of Hawai'i Part 2, Spatial and Temporal Variations in Salinity, Radium-Isotope Activity, and Nutrient Concentrations in Coastal Waters, December 2003-April 2006. USGS Scientific Investigations Report 2008-5128.

Knee, K.L., R.L. Leopold, E.R. Madsen, A. Paytan.2008. Assessing the importance of sand as a source of fecal indicator bacteria. Oceanography 21(3):72-80.

Knee,K.L., B.A. Layton, J.H. Street, A.B. Boehm, and A. Paytan. 2008. Sources of nutrients and fecal indicator bacteria to nearshore waters on the north shore of Kaua`i (Hawai`i, USA). Estuaries and Coasts doi. 10.1007/s12237-008-9055-6.

Lazar, B., Y. Weinstein, A. Paytan, E. Magal, D. Bruce, and Y. Kolodny. 2008. Ra and Th adsorption coefficients in lakes -- Lake Kineret (Sea of Galilee) "natural experiment". Geochimica et Cosmochimica Acta 72: 3446-3459.

Mackey, K. R. M., A. Paytan, A.R. Grossman and S. Bailey. 2008. A photosynthetic strategy for coping in a high-light, low-nutrient environment. Limnology and Oceanography 53(3): 900-913.

Chen, Y., A. Paytan, Z. Chase, C. Measures, A.J. Beck, S.A. Sañudo-Wilhelmy, and A.F. Post. 2008. Sources and fluxes of atmospheric trace elements to the Gulf of Aqaba, Red Sea. Journal of Geophysical Research 113: D05306.

Griffith, E.M., A. Paytan, R. Kozdon, A. Eisenhauer, A. C. Ravelo. 2008. Influences on the fractionation of calcium isotopes in planktonic foraminifera. Earth and Planetary Science Letters 268: 124-136.

Geibert, W., M. Charette, G. Kim, W.S. Moore, J. Street, M. Young, and A. Paytan. 2008. The release of dissolved actinium to the ocean: A global comparison of different end-members. Mar. Chem. 109(3-4): 409-420.

Street, J.H., K.L. Knee, E.E. Grossman, and A. Paytan. 2008. Submarine groundwater discharge and nutrient addition to the coastal zone and coral reefs of leeward Hawai`i. Mar. Chem. 109(3-4): 355-376.

Young, M.B., M.E. Gonneea, D.A. Fong, W.S. Moore, J. Herrera-Silveira, and A. Paytan. 2008. Characterizing sources of groundwater to a tropical coastal lagoon in a karstic area using radium isotopes and water chemistry. Mar. Chem. 109(3-4): 377-394.


Feranec, R.S., E.A. Hadley, J.L. Blois, A.D. Barnosky, and A. Paytan. 2007. Radiocarbon dates from the Pleistocene fossil deposits of Samwel Cave, Shasta County, California, USA. Radiocarbon 49(1): 117-121.

Paytan, A., K. Averyt, K. Faul, E. Gray, and E. Thomas. 2007. Barite accumulation, ocean productivity, and Sr/Ba in barite across the Paleocene-Eocene Thermal Maximum. Geology 35(12): 1139-1142.

Saltzman, J. and A. Paytan. 2007. Engaging first-grade students in a geoscience campus field trip. Journal of Geoscience Education 55(4): 326-332.

Feranec, R. S., E. A. Hadley and A. Paytan. 2007. Determining landscape use of Holocene mammals using strontium isotopes. Oecologia 153: 943-950.

Derse, E., K. Knee, S. Wankel, C. Kendall, C.J. Berg, Jr. and A. Paytan. 2007. Identifying the sources of nitrogen to Hanalei Bay, Kaua'i utilizing the nitrogen isotope signature of macroalgae. Environ. Sci. Technol. 41:5217-5223.

Paytan, A. and K. McLaughlin. 2007. Phosphorus in Our Waters. Oceanography 20(2): 200-206.

Leichter, J.J., A. Paytan, S. Wankel, K. Hanson, S. Miller and M.A. Altabet. 2007. Nitrogen and oxygen isotopic signatures of subsurface nitrate: evidence of deep water nutrient sources to the Florida Keys reef tract. Limnology and Oceanography 52(3) 1258-1267.

Paytan, A. and E. M. Griffith. 2007. Marine Barite: Recorder of variations in ocean export productivity. Deep-Sea Research II: 687-705.

Wankel, S., C. Kendall, T. Pennington, F. Chavez, and A. Paytan. 2007. Nitrification in the euphotic zone as evidenced by nitrate dual isotopic composition: Observations from Monterey Bay, California. Global Biogeochemical Cycles 21, doi:10.1029/2006GB002723

Paytan, A. 2007. Journal Club. Nature 446: 587.

Vengosh, A., S. Hening, J. Ganor, B. Mayer, C.E. Weyhenmeyer, T.D. Bullen and A. Paytan. 2007. New isotopic evidence for the origin of groundwater from the Nubian Sandstone Aquifer in the Negev, Israel. Applied Geochemistry. doi:10.1016/j.apgeochem.2007.01.005.

Mackey, K.R.M., R.G. Labiosa, M. Calhoun, J.H. Street, A. Post and A. Paytan. 2007. Phosphorus availability, phytoplankton community dynamics, and taxon specific phosphorus status in the Gulf of Aqaba, Red Sea. Limnology and Oceanography 52:873-885.

Chen, Y., S. Mills, J. Street, D. Golan, A. Post, M. Jacobson and A. Paytan. 2007. Estimates of atmospheric dry deposition and associated input of nutrients to Gulf of Aqaba seawater. Journal of Geophysical Research 112, D04309,doi:10.1029/2006JD007858.

Gallego-Torres, D., F. Martinez-Ruiz, A. Paytan, F.J. Jiménez-Espejo and M. Ortega-Huertas. 2007. Pliocene-Holocene evolution of depositional conditions in the eastern Mediterranean: Role of anoxia vs. productivity at time of sapropel deposition. Palaeogeography, Palaeoclimatology, Palaeoecology doi:10.1016/j.palaeo.2006.10.008.

Lyle, M., N. Pisias, A. Paytan, J. I. Martinez and A. Mix. 2007. Reply to comment by R. Francois et al. on "Do geochemical estimates of sediment focusing pass the sediment test in the equatorial Pacific?": Further explorations of 230Th normalization. Paleoceanography 22: doi: 10.1029/2006PA001373.

Paytan, A. and M.L. Zoback. 2007. The inside track from academia and industry: Crossing boundaries, hitting barriers. Nature 445: 950.

Jiménez-Espejo, F.J., F. Martínez-Ruiz, A. Paytan, T. Sakamoto, M. Ortega-Huertas, G. Finlayson, K. Iijima, D. Gallego-Torres and D. Fa. 2007. Climate forcing and Neanderthal extinction in Southern Iberia: insights from a multiproxy marine record. Quaternary Science Reviews doi: 10.1016/j.quascirev.2006.12.013.

Paytan, A. and K. McLaughlin. 2007. The Oceanic Phosphorus Cycle. Chemical Reviews 107:563-576.

Averyt, K.B. and A. Paytan. 2007. Reply to Comment on “Empirical partition coefficients for Sr and Ca in marine barite: Implications for reconstructing seawater Sr and Ca concentrations”. Geochemistry, Geophysics, Geosystems 8: Q01009, doi:10.1029/2006GC001494.


McLaughlin, K., B.J. Cade-Menun and A. Paytan. 2006. The oxygen isotopic composition of phosphate in Elkhorn Slough, California: A tracer for phosphate sources. Estuarine, Coastal and Shelf Science 70: 499-506.

McLaughlin, K., F. Chavez, J.T. Pennington and A. Paytan. 2006. A time series investigation of the oxygen isotopic composition of dissolved inorganic phosphate in Monterey Bay, California. Limnology and Oceanography 51:2370-2379.

Handler, N.B., A.Paytan, C.P. Higgins, R.G. Luthy and A.B. Boehm. 2006. Human development is linked to multiple water body impairments along the California coast. Estuaries and Coasts 29: 860-870.

Chase, Z., A. Paytan, K.S. Johnson, J. Street and Y. Chen. 2006. Input and cycling of iron in the Gulf of Aqaba, Red Sea. Global Biogeochem. Cycles 20: GB3017, doi: 10.1029/2005GB002646.

Chen, Y., J. Street and A. Paytan. 2006. Comparison between pure-water- and seawater-soluble nutrient concentrations of aerosols from the Gulf of Aqaba. Mar. Chem. 101: 141-152.

Böhm, F., N. Gussone, A. Eisenhauer, W-C. Dullo, S. Reynaud and A. Paytan. 2006. Calcium isotope fractionation in Modern scleractinian corals. Geochimica et Cosmochimica Acta 70: 4452-4462.

Hoppe, K.A., A. Paytan and P. Chamberlain. 2006. Reconstructing Grassland vegetation and paleotemperatures using carbon isotope ratios of bison tooth enamel. Geology 34: 649-652.

Wei, Z., J.M. Moldowan and A. Paytan. 2006. Diamondoids and molecular biomarkers generated from modern sediments in the absence and presence of minerals during hydrous pyrolysis. Organic Geochemistry 37: 891-911.

McLaughlin, K., C. Kendall, S.R. Silva, M. Young and A. Paytan. 2006. Phosphate oxygen isotope ratios as a tracer for sources and cycling of phosphate in North San Francisco Bay, California. Journal of Geophysical Research 111: G03003.

Wankel, S.D., C. Kendall, C.A. Francis and A. Paytan. 2006. Nitrogen sources and cycling in the San Francisco Bay Estuary: A nitrate dual isotopic composition approach. Limnology and Oceanography 51: 1654-1664.

Shellenbarger, G.G., S.G. Monismith, A. Genin and A. Paytan. 2006. The importance of submarine groundwater discharge to the nearshore nutrient supply in the Gulf of Aqaba (Israel). Limnology and Oceanography 51: 1876-1886.

Nicholson, D., S. Dyhrman, F. Chavez and A. Paytan. 2006. Alkaline phosphatase activity in the Phytoplankton communities of Monterey Bay and San Francisco Bay. Limnology and Oceanography 51: 874-883.

Gonneea, M. E. and A. Paytan. 2006. Phase associations of barium in marine sediments. Mar. Chem. 100:124-135.

Boehm, A.B., A. Paytan, G.G. Shellenbarger and K.A. Davis. 2006. Composition and flux of groundwater from a California beach aquifer: Implications for nutrient supply to the surf zone. Continental Shelf Research 26: 269-282.

McLaughlin, K., A. Paytan, C. Kendall and S. Silva. 2006. Oxygen isotopes of phosphatic compounds—Application for marine particulate matter, sediments and soils. Marine Chemistry 98:148-155.

Paytan, A. , G.G. Shellenbarger, J.H. Street, M.E. Gonneea, K. Davis, M.B. Young and W.S. Moore. 2006. Submarine groundwater discharge: An important source of new inorganic nitrogen to coral reef ecosystems. Limnology and Oceanography 51: 343-348.


Monismith, S., Jones, N., Bela, M., NidziekoDe. N., A. Paytan , Misra, G. and Street, J. 2005. Hydrodynamics and sediment dynamics in Elkhorn Slough: A report to Monterey Bay Sanctuary Foundation

De La Rocha, C.L. and A. Paytan. 2005. Editorial: Directions in the study of ocean chemistry over The Phanerozoic and its links to geologic processes. Marine Geology 217: 193-197.

Young, M., M.E. Gonneea, J. Herrera-Silveira and A. Paytan. 2005. Export of dissolved and particulate carbon and nitrogen from a mangrove-dominated lagoon, Yucatan Peninsula, Mexico. International Journal of Ecology and Environmental Sciences 31: 189-202.

Cade-Menun, B.J., C.R. Benitez-Nelson, P. Pellechia and A. Paytan. 2005. Refining 31P nuclear magnetic resonance spectroscopy for marine particulate samples: Storage conditions and extraction recovery. Mar. Chem. 97: 293-306.

Faul, K. L., A. Paytan, and M. L. Delaney. 2005. Phosphorus distribution in sinking organic particulate matter. Mar. Chem. 97: 307-333.

Faul, K. and A. Paytan. 2005. Phosphorus and barite concentrations and geochemistry in Site 1221 Paleocene/Eocene boundary sediments. Proceedings of the Ocean Drilling Program, Scientific Results Volume 199.

Averyt, K.B., M. Calhoun, L. Schmalz and A. Paytan. 2005. Data report: Carbonate and barite trends across the Eocene/Oligocene boundary at Shatsky Rise, ODP Leg 198. Proceedings of the Ocean Drilling Program, Scientific Results Volume 198.

Porder, S., A. Paytan and P.M. Vitousek. 2005. Erosion and landscape development affect plant nutrient status In the Hawaiian Islands. Oecologia 142: 440-449.

Lyle, M., N. Mitchell, N. Pisias, A. Mix, J.I. Martinez and A. Paytan. 2005. Do geochemical estimates of sediment Focusing pass the sediment test in the equatorial Pacific? Paleoceanography 20: doi: 10.1029/2004PA001019.

Street, J. and A. Paytan. Iron, phytoplankton growth, and the carbon cycle. Biogeochemical Cycles of elements. Volume 43 of Metal ions in biological systems. Eds. Astrid Sigel, Helmut Sigel and Roland R.O. Sigel. Boca Raton: Taylor and Francis. 2005. 153-185.


Paytan, A., F. Martinez-Ruiz, M. Eagle, A. Ivy, and S.D. Wankel. 2004. Using sulfur isotopes to elucidate the origin of barite associated with high organic matter accumulation events in marine sediments. Sulfur Biogeochemistry, GSA Special Paper, 379, 151-160.

Paytan, A. 2004. Biogeochemistry of Marine Systems, (Ed. Kenneth D. Black and Graham B. Shimmield) Blackwell Publishing, book review, Oceanography 17, 213-214.

Paytan, A., M. Lyle, A. Mix, and Z. Chase. 2004. Climatically Driven changes in oceanic processes throughout the equatorial Pacific. Paleoceanography 19: 4017-4023.

Averyt, K.B. andA. Paytan. 2004. A comparison of multiple proxies for export production in the equatorial Pacific. Paleoceanography 19: 4003-4016.

Gonneea, M.E., A. Paytan, and J.A. Herrera-Silviera. 2004. Tracing Organic matter sources and carbon burial in mangrove sediments over the past 160 years. Estuarine, Coastal and Shelf Science 61: 211-227.

McLaughlin K., S. Silva, C. Kendall, H. Stuart-Williams and A. Paytan. 2004. A precise method for the analysis of face="Symbol">d18O of dissolved inorganic phosphate in seawater. Limnology and Oceanography: Methods 2: 202-212.

Paytan, A., A.B. Boehm, and G.G. Shellenbarger. 2004. Bacterial contamination and submarine groundwater discharge - a possible link. Environmental Chemistry 1, 1-2, doi: 10.1071/EN04002.

Paytan, A., M. Kastner, D. Campbell and M.H. Thiemens. 2004. Seawater sulfur isotope fluctuations in the Cretaceous. Science 304: 1663-1665.

Boehm, A. B., G.G. Shellenbarger and A. Paytan. 2004. Groundwater discharge: Potential association with fecal indicator bacteria in the surf zone. Environmental Science and Technology 38: 3558-3566.


Kurtz, A.C., L.R. Kump, M.A. Arthur, J.C. Zachos and A. Paytan. 2003. Early Cenozoic decoupling of the global carbon and sulfur cycles. Paleoceanography 18: 1090, 10.1029/2003PA000908.

González-Muñoz, M.T., B. Fern á ndez-Luque, F. Martínez-Ruiz, K.B. Chekroun, J. M. Arias, M. Rodr í guez-Gallego, M. Mart í nez-Ca ñ amero, C. de Linares and A. Paytan. 2003. Precipitation of barite by yxococcus xanthus: Possible Implications for the Biogeochemical Cycle of Barium. Applied and Environmental Microbiology 69: 5722-5725.

Paytan, A., B. Cade-Menun, K. McLaughlin and K. L. Faul. 2003. Selective phosphorus regeneration of sinking marine particles: evidence from 31P-NMR, Marine Chemistry 82: 55-70.

Averyt, K.B. and A. Paytan. 2003. Empirical partition coefficients for Sr and Ca in marine barite: Implications for reconstructing seawater Sr and Ca concentrations. G3 4: 1043.

Averyt, K.B., A. Paytan and G. Li. 2003. A precise, high-throughput method for determining Sr/Ca, Sr/Ba, and Ca/Ba ratios of marine barite. G3 4: 1039.

Porder, S. A. Paytan and E.A. Hadly. 2003. Mapping the origin of faunal assemblages using strontium isotopes. Paleobiology 29: 197-204.

Martínez-Ruiz, F., A. Paytan, M. Kastner, J.M. González-Donoso, D. Linares, S.M. Bernasconi and F.J. Jimenez-Espejo. 2003. A comparative study of the geochemical and mineralogical characteristics of the S1 sapropel in the western and eastern Mediterranean. Palaeo. Palaeo. Palaeo. 190: 23-37.

Eagle, M., A. Paytan, K.R. Arrigo, G. van Dijken, and R.W. Murray. 2003. A comparison between excess barium and barite as indicators of carbon export. Paleoceanography 18. doi: 10.1029/2002PA000793. **Please note: The units of Ba Excess AR and barite AR in Table 3 and Figure 3 should be mg/m2y

Mearon, S., A. Paytan, and T.J. Bralower. 2003. Cretaceous Strontium isotope stratigraphy using marine barite. Geology 31: 15-18.

Page last updated on June 18, 2018